Vannellina unifies all
fan-shaped, semicircular or spatulate amoebae of vannellian
morphotypes (the genera Vannella, Platyamoeba, and
Clydonella), as well as the genus Lingulamoeba,
which is more lingulate than fan-shaped. The existence of
this clade was first shown by Peglar et al. (2003), and it
corresponds to the family Vannellidae in the traditional
classification (Page 1987).
The synapomorphic characters of
the clade are the flattened, generally fan-shaped
locomotive form without pseudopodia, and the rolling
movement of the cell membrane during locomotion.