Nicolas Ray Group

Nicolas Ray - full list of publications

Citations of my publications are found on Google Scholar Citation

International peer reviewed publications (articles and book chapters)

  1. Branco C, Ray N, Currat M & M Arenas. 2020. Influence of Paleolithic range contraction, admixture and long-distance dispersal on genetic gradients of modern Humans in Asia. Molecular Ecology, 29: 2150-2159 [link to article]
  2. Hierink F, Bolon I, Durso AM, Ruiz de Castañeda R, Zambrana-Torrelio C, Eskew E & N Ray. 2020. Forty-four years of global trade in CITES-listed snakes: trends and implications for conservation and public health. Biological Conservation, 248: 108601 [link to article]
  3. Ochoa C, Bolon I, Ruiz de Castañeda R, Alcoba G, Durso A, Babo Martins S, Chappuis F & N Ray. 2020. Assessing the increase of snakebite incidence in relationship to flooding events. Journal of Environmental and Public Health, 6135149 [link to article]
  4. Bolon I, Durso AM, Botero Mesa S, Ray N, Alcoba G, Chappuis F & R Ruiz de Castañeda. 2020. Identifying the snake: first scoping review on practices of communities and healthcare providers confronted with snakebite across the world. PLoS One, 15(3): e0229989 [link to article]
  5. Ruiz de Castañeda R, Sheath D, Bempong N, Raviglione M, Machalaba C, Pepper M.S., Vayena E, Ray N, Wernli D, Escher G, Grey F, Elger B, Helbing D, Kleineberg K, Beran DH, Miranda J, Huffman MD, Hersch F, Andayi F, Thumbi SL, D'Acremont V, Hartley M-A, Zinsstag J, Larus J, Rodriguez M, Guerin P, Merson L, Ngyuen V-K, Rhuli F, Geissbuhler A, Salathé M, Bolon I, Boheme CR, Berkley SR, Valleron A-J, Keiser O, Kaiser L, Eckerle I, Utzinger J & A Flahault. 2020. Precision Global Health: a roadmap for augmented action. Journal of Public Health and Emergency, 4: 6 [link to article]
  6. Miranda Espinosa MT, Giuliani G & N Ray. 2020. Reviewing the discoverability and accessibility to data and information products linked to Essential Climate Variables. International Journal of Digital Earth, 13(2): 236-252 [link to article]
  7. Lehmann A, Nativi S, Mazzetti P, Maso J, Serral I, Spengler D, Niamir A, McCallum I, Lacroix P, Patias P, Rodila D, Ray N & G Giuliani. 2020. GEOEssential - mainstreaming workflows from data sources to environment policy indicators with essential variables. International Journal of Digital Earth, 13(2): 322-338 [link to article]
  8. Ranzoni J, Giuliani G, Huber L & N Ray. 2019. Modelling the nocturnal ecological continuum of the State of Geneva, Switzerland, based on high-resolution nighttime imagery. Remote Sensing Applications: Society and Environment, 16: 100268 [link to article
  9. Ruiz de Castañeda R, Durso AM, Ray N, Fernández JL, Williams D, Alcoba G, Chappuis F, Salathé M & I Bolon. 2019. Snakebite and snake identification: empowering neglected communities and health-care providers with AI. The Lancet Digital Health, 1(5): e202-e203 [link to article]
  10. Babo Martins S, Bolon I, Chappuis F, Ray N, Alcoba G, Ochoa C, Sharma SJ, Nkwescheu A, Wanda F, Durso AM & R Ruiz de Castañeda. 2019. Snakebite and its impact in poor rural communities: The need for a One Health approach. PLoS Neglected Tropical Diseases, 13(9): e0007608 [link to article]
  11. Ross N, Eskew EA & N Ray. 2019. Citesdb: An R package to support analysis of CITES Trade Database shipment-level data. Journal of Open Source Software, 4(37): 1483 [link to article]
  12. Currat M, Arenas M, Quilodran C, Excoffier L & N Ray. 2019. SPLATCHE3: simulation of serial genetic data under spatially explicit evolutionary scenarios including long-distance dispersal. Bioinformatics, 35(21): 4480-4483 [link to article]
  13. Kogan NE, Bolon I, Ray N, Alcoba G, Fernandez-Marquez JL, Müller MM, Mohanty SP & R Ruiz de Castañeda. 2019. Wet markets and food safety: TripAdvisor for improved global digital surveillance? JMIR Public Health & Surveillance 5(2): e11477 [link to article]
  14. Ebener S, Stenberg K, Brun M, Monet J-P, Ray N, Sobel H, Roos N, Gault P, Morrisey Conlon C, Bailey P, Moran AC, Ouedraogo L, Kitong J, Ko E, Sanon D, Jega FM, Azogu O, Ouedraogo B, Osakwe C, Chimwemwe Chanza H, Steffen M, Ben Hamadi I, Tib H, Haj Asaad A & T Tan Torres. 2019. Proposing standardised geographical indicators of physical access to Emergency Obstetric and Newborn Care in low- and middle-income countries. BMJ Global Health 4: e000778 [link to article]
  15. Lacroix P, Moser F, Benvenuti A, Piller T, Jensen D, Petersen I, Planque M & N Ray. 2019. MapX: an open geospatial platform to manage, analyse and visualize data on natural resources and the environment. SoftwareX 9: 77-84 [link to article]
  16. Branco C, Velasco M, Benguigui M, Currat M, Ray N & M Arenas. 2018. Consequences of diverse evolutionary processes on American genetic gradients of modern humans. Heredity  121: 548-556 [link to article]
  17. Longbottom J, Shearer FM, Devine M, Alcoba G, Chappuis F, Weiss DJ, Ray S, Ray N, Warrell D, De Castañeda RR, Williams DJ, Hay SI & D Pigott. 2018. Vulnerability to snakebite envenoming: a global mapping of hotspots. The Lancet 392(10148): 673-684 [link to article]
  18. Geneviève L, Ray N, Chappuis F, Alcoba G, Mondardini R, Bolon I & R Ruiz de Castañeda. 2018. Participatory approaches and open data on venomous snakes: A neglected opportunity in the global snakebite crisis? PLoS Neglected Tropical Diseases 12(3), e0006162 [link to article]
  19. Atibu EK, Lacroix P, Sivalingam P, Ray N, Giuliani G, Mulaji CK, Otamonga J-P, Mpiana PT, Slaveykova VI & J Poté. 2018. Persistent high contamination in the areas surrounding abandoned mines and mining activities: an impact assessment of the Dilala, Luilu and Mpingiri Rivers, Democratic Republic of the Congo. Chemosphere 191: 1008-1020 [link to article]
  20. Yushchenko A, de Bono A, Chatenoux B, Patel MK & N Ray. 2018. GIS-based assessment of photovoltaic (PV) and concentrated solar power (CSP) generation potential in West Africa. Renewable and Sustainable Energy Reviews 81(2): 2088-2103 [link to article]
  21. Lehmann A, Guigoz Y, Ray N, Mancuso E, Abbaspour K, Rouholahnejad E, Allenbach K, De Bono A,  Fasel M, Gago-Silva A, Bär R, Lacroix P. & G Giuliani. 2017. A regional platform from past to future land uses, climate, demography, hydrology and beaches in the Black Sea catchment. Nature - Scientific Data 4: 170087 [link to article]
  22. Gago da Silva A, Ray N & Lehmann A. 2017. Spatial dynamic modeling of future scenarios of land use change in Vaud and Valais, Western Switzerland. International Journal of Geo-Information, 6: 115 [link to pdf]
  23. Desconnets JC, Giuliani G, Guigoz, Y, Lacroix P, Mlisa A, Noort M, Ray N & ND Searby. 2017. GEOCAB Portal: a gateway for discovering and accessing capacity building resources in Earth Observation. International Journal of Applied Earth Observation and Geoinformation 54:95-104 [link to article]
  24. Kasparian J, Hassler C, Ibelings B, Berti N, Bigorre S, Djambazova V, Gascon-Diez E, Giuliani G, Houlman R, Kiselev D, de Laborie P, Le AD, Magouroux T, Neri T, Palomino D, Pfänder S, Ray N, Sousa G, Stadler D, Tettamanti F, Wolf J-P & M Beniston. 2017. Assessing the Dynamics of Organic Aerosols over the North Atlantic Ocean. Nature - Scientific Reports 7: 45476  [link to article]
  25. Amorim CEG, Hofer T, Ray N, Foll M, Ruiz-Linares A & L Excoffier. 2017. Long-distance dispersal suppresses introgression of local alleles during range expansions. Heredity 118(2): 135-142 [link to article]
  26. Guigoz Y, Lacroix P, Rouholahnejad E, Ray N & G Giuliani. 2017. SCOPED-W: SCalable Online Platform for extracting Environmental Data and Water-related model outputs. Transactions in GIS 21(4): 748-763 [link to article]
  27. Fontaine MC, Thatcher O, Ray N, Piry S, Brownlow A, Davison NJ, Jepson P, Deaville R & SJ Goodman. 2017. Mixing of Porpoise ecotypes in South Western UK waters revealed by genetic profiling. Royal Society Open Science 4(3): 160992 [link to article]
  28. Rahman K, Ray N, Giuliani G, Schwank J, Girón G, Escobar R, George C & A Lehmann. 2017. Breaking walls towards fully open source hydrological modeling. Water Resources 44(1), 23-30 [link to article]
  29. Mazzetti P, Roncella R, Mihon D, Bacu V, Lacroix P, Guigoz Y, Ray N, Giuliani G, Gorgan D & S Nativi. 2016. Integration of data and computing infrastructures for Earth Science: an image mosaicking use-case. Earth Science Informatics 9(3): 325-342 [link to article]
  30. Giuliani G, Lacroix P, Guigoz Y, Roncella R, Bigagli L, Santoro M, Mazzetti P, Nativi S, Ray N & A. Lehmann. 2017. Bringing GEOSS services into practice: a capacity building resource on spatial data infrastructures (SDI). Transactions in GIS 21(4): 811-824 [link to article]
  31. Guigoz Y, Giuliani G, Nonguierma A, Lehmann A, Mlisa A & N Ray. 2017. Spatial Data Infrastructure in Africa: a gap analysis. Journal of Environmental Informatics 21(4): 748-763 [link to article]
  32. Ray N, Lacroix P, Giuliani G, Upla P, Rajabifard A & D Jensen. 2016. Open Spatial Data Infrastructures for the Sustainable Development of the Extractives Sector: Promises and Challenges. Ed. Coleman D, Rajabifard A, Crompvoets J. In: "Spatial Enablement in a Smart World". GSDI Association press, 53-69  [pdf]
  33. Giuliani G, Guigoz Y, Lacroix P, Ray N & A Lehmann. 2016. Facilitating the production of ISO-compliant metadata of geospatial datasets. International Journal of Applied Earth Observation and Geoinformation 44: 239-243 [pdf]
  34. Alves I, Arenas M, Currat M, Sramkova Hanulova A, Sousa V, Ray N & L Excoffier. 2016. Long distance dispersal shaped patterns of human genetic diversity in Eurasia. Molecular Biology and Evolution 33(4):946–958 [pdf][suppl. mat.]
  35. Rodila D, Ray N & D Gorgan. 2015. Conceptual model for environmental science applications on parallel and distributed infrastructures. Environmental Systems Research 4:23 [pdf]
  36. Myroshnychenko V, Ray N, Lehmann A, Giuliani G, Kideys A, Weller P & Teodor D. 2015. Environmental data gaps in Black Sea catchment countries: INSPIRE and GEOSS State of Play. Environmental Science and Policy 46:13-25 [pdf]
  37. Lehmann A, Giuliani G, Mancuso E, Abbaspour KC, Sozen S, Gorgan D, Beel A & Ray N. 2015. Filling the gap between Earth observation and policy making in the Black Sea catchment with enviroGRIDS. Environmental Science and Policy 46:1-12 [pdf]
  38. Astsatryan H, Hayrapetyan A, Narsisian W, Saribekyan A, Asmaryan Sh, Muradyan V, Guigoz Y, Giuliani G & Ray N. 2015. An interoperable web portal for parallel geoprocessing of satellite image vegetation indices. Earth Science Informatics 8(2): 453-460 [pdf]
  39. Astsatryan H, Hayrapetyan A, Narsisian W, Asmaryan S, Saghatelyan A, Muradyan V, Guigoz Y, Giuliani G & Ray N. 2015. An Interoperable Cloud-based scientific gateway for NDVI time series analysis. Computer Standards & Interfaces 41, 79-84 [pdf]
  40. Asmaryan S, Saghatelyan A, Astsatryan H, Bigagli L, Mazzetti P, Nativi S, Guigoz Y, Lacroix P, Giuliani G & N Ray. 2014. Leading the way toward an environmental National Spatial Data Infrastructure in Armenia. South-Eastern European Journal of Earth Observation and Geomatics 3: 53-62 [pdf]
  41. Ruiz-Linares A, Adhikari K, Acuña-Alonzo V, Quinto M, Jaramillo C, Arias W, Fuentes M, Pizarro M, Everardo P, de Avila F, Gómez-Valdés J, León-Mimila P, Hunemeier T, Ramallo V, Silva de Cerqueira CC, Burley M-W, Konca E, Zagonel de Oliveira M, Roberto Veronez M, Rubio-Codina M, Attanasio O, Gibbon S, Ray N, Gallo C, Poletti G, Rosique J, Schuler-Faccini L, Salzano FM, Bortolini M-C, Canizales-Quinteros S, Rothhammer F, Bedoya G, Balding D & Gonzalez-José R. 2014. Admixture in Latin America: geographic structure, phenotypic diversity and self-perception of ancestry based on 7,342 individuals. PLoS Genetics 10(9):e1004572 [pdf]
  42. Constantinescu A, Hanganu J, Lehmann A & N Ray. 2014. Predicting the spatial shifts of alpine habitats from climate change scenarios in Romania. Romanian Journal of Geography 58(2): 115-127 [pdf]
  43. Lehmann A, Giuliani G, Ray N, Rahman K, Abbaspour KC, Nativi S, Craglia M, Cripe D, Quevauviller P & Beniston M. 2014. Reviewing innovative Earth Observation solutions for filling science-policy gaps in hydrology. Journal of Hydrology 518(B):267-277 [pdf]
  44. Ray N. 2013. Les origines de l'humanité: l'apport de la génétique. In: "Adam, qui es-tu? Perspectives bibliques et scientifiques sur l'origine de l'humanité". Ed., Jaeger, L. Editions Excelsis, Charols, 89-109 [link to book]
  45. Cau P, Manca S, Soru C, Muroni D, Gorgan D, Bacu V, Lehmann A, Ray N & Giuliani G. 2013. An interoperable OGC-compliant Decision Support System for Water Resources Management. International Journal of Advanced Computer Science and Applications 3(3):75-82 [pdf]
  46. Gorgan D, Giuliani G, Ray N, Lehmann A, Cau P, Abbaspour K, Charvat K & Jonoski A. 2013. Black Sea Catchment Observation System as a Portal for GEOSS Community. International Journal of Advanced Computer Science and Applications 3(3):9-18 [pdf]
  47. Giuliani G, Rahman K, Ray N & Lehmann A. 2013. OWS4SWAT: Publishing and Sharing SWAT Outputs with OGC standards. International Journal of Advanced Computer Science and Applications 3(3):90-98 [pdf]
  48. Giuliani G, Ray N & Lehmann A. 2013. Building Regional Capacities for GEOSS and INSPIRE: a journey in the Black Sea Catchment. International Journal of Advanced Computer Science and Applications 3(3):19-27 [pdf]
  49. Arenas M, Mona S, Trochet A, Sramkova Hanulova A, Currat M, Ray N, Chikhi L, Rasteiro R, Schmeller DS & Excoffier L. 2014. The scaling of genetic diversity in a changing and fragmented world. In: Scaling in ecology and conservation, eds. Henle et al. (Pensoft Publishers), pp. 55-60 [pdf]
  50. Lacroix P, de Roulet P & Ray N. 2014. Simplified Toolbar to Accelerate Repeated Tasks (START) for ArcGIS: optimizing workflows in humanitarian demining. International Journal of Applied Geospatial Research 5(4): 87-94 [link to pdf]
  51. Geiser C, Ray N, Lehmann A & Ursenbacher S. 2013. Unravelling landscape variables with multiple approaches to overcome scarce species knowledge: A landscape genetic study of the slow worm. Conservation Genetics 14(4): 783-794 [link to pdf]
  52. Lacroix P, Santiago H & Ray N. 2014. MASCOT: Multi-criteria Analytical SCOring Tool for ArcGIS Desktop. International Journal of Information, Technology and Decision Making 13(6): 1135-1159 [pdf]
  53. Mona S, Ray N, Arenas M & Excoffier L. 2014. Genetic consequences of habitat fragmentation during a range expansion. Heredity 112: 291-299 [pdf]
  54. Yalew SG, van Griensven A, Ray N, Kokoszkiewicz L. & Betrie GD. 2013. Distributed computation of large scale SWAT models on the Grid. Environmental Modelling & Software 41: 223-230 [pdf]
  55. Arenas M, François O, Currat M, Ray N & Excoffier L. 2013. Influence of admixture and Paleolithic range contractions on current European diversity gradients. Molecular Biology and Evolution 30(1): 57-61 [pdf] [suppl. mat.]
  56. Reich D, Patterson N, Campbell D, Tandon A, Mazieres S, Ray N, Parra MV, Rojas W, DuqueMesa, N, Velez ID, García LF, Triana O, Blair S, Maestre A, Dib JC, Bravi CM, Bailliet G, Corach D, Hünemeier T, Bortolini MC, Salzano F, Petzl-Erler ML, Acuña-Alonzo V, Canizales-Quinteros S, Aguilar-Salinas C, Tusié-Luna T, Riba L, Rodríguez-Cruz M, Lopez-Alarcón M, Coral-Vazquez R, Canto-Cetina T, Silva- Zolezzi I, Fernandez-Lopez JC, Contreras AV, Jimenez-Sanchez G, Gómez-Vázquez MJ, Molina J, Carracedo A, Salas A, Gallo C, Poletti G, Witonsky DB, Alkorta-Aranburu G, Sukernik R, Osipova L, Fedorova S, Vasquez R, Villena M, Moreau C, Hammer M, Barrantes R, Pauls D, Excoffier L, Bedoya G, Rothhammer F, Dugoujon JM, Larrouy G, Klitz W, Labuda D, Kidd J, Kidd K, Di Rienzo A, Freimer NB, Price AL & Ruiz-Linares A. 2012. Reconstructing native American population history. Nature 488: 370-374 [Link to article] [Press release from UNIGE] [NY Times article] [LA Times article] [BBC article]
  57. Gorgan D, Bacu V, Mihon D, Rodila D, Stefanut T, Abbaspour K, Cau P, Giuliani G, Ray N & Lehmann A. 2012. Spatial data processing tools and applications for Black Sea catchment region. International Journal of Computing 11(4): 336-344 [pdf]
  58. Gorgan D, Bacu V, Mihon D, Stefanut T, Rodila D, Cau P, Abbaspour K, Giuliani G, Ray N & Lehmann A. 2012. Software platform interoperability throughout enviroGRIDS portal. Journal of Selected Topics in Applied Earth Observations and Remote Sensing 5(6): 1617-1627 [pdf]
  59. Kormann U, Gugerli F, Ray N, Excoffier L & Bollmann K. 2012. Parsimony-based pedigree analysis and resistance models suggest topography to restrict dispersal and connectivity in the endangered capercaillie. Biological Conservation 52: 241-252 [Link to pdf and suppl. mat.]
  60. Arenas M, Ray N, Currat M & Excoffier L. 2012. Consequences of range contractions and range shifts on molecular diversity. Molecular Biology and Evolution 29(1): 207-218 [pdf] [suppl. mat.]
  61. Gorgan D, Abbaspour K, Cau P, Bacu V, Mihon D, Giuliani G, Ray N & Lehmann A. 2011. Grid based data processing tools and applications for black sea catchment basin. Proceedings of IDAACS (1) 2011: 223-228 [pdf]
  62. Ray N, Giuliani G, Gorgan D & Lehmann A. 2011. Distributed geocomputation for modeling the hydrology of the Black Sea watershed. Watershed Approach to Environmental Security: Fostering integrated water management in the Azov Sea Basin, ed. Lagutov, V. NATO Science for Peace and Security Series: Environmental Security. Dordrecht, Springer. Pp. 141-158
  63. Giuliani G, Nativi S, Lehmann A & Ray N. 2012. WPS mediation: an approach to process geospatial data on different computing backends. Computers and Geosciences 47: 20-33 [pdf]
  64. Lecca G, Petitdidier M, Hluchy L, Ivanovic M, Kussul N, Ray N & Thieron V. 2011. Grid computing technology for hydrological applications. Journal of Hydrology 403: 186-199 [pdf]
  65. Giuliani G, Ray N, Schwarzer S, de Bono A, Dao H, Peduzzi P, Beniston M, Van Woerden J, Witt R & Lehmann A. 2011. Sharing environmental data through GEOSS. International Journal of Applied Geospatial Research 2(1): 1-17 [pdf]
  66. Giuliani G, Ray N & Lehmann A. 2011. Grid-enabled Spatial Data Infrastructure for environmental sciences: challenges and opportunities. Future Generation Computer Systems 27(3): 292-303 [pdf]
  67. Ray N, Currat M, Foll M. & Excoffier L. 2010. SPLATCHE2: a spatially-explicit simulation framework for complex demography, genetic admixture and recombination. Bioinformatics 26(23): 2993-2994 [pdf]
  68. Yang NN, Mazières S, Bravi C, Ray N, Wang S, Burley M-W, Bedoya BG, Rojas W, Parra MV, Molina JA, Gallo C, Poletti G, Hill K, Hurtado AM, Petzl-Erler ML, Tsuneto LT, Klitz W, Barrantes R, Llop E, Rothhammer F, Labuda D, Salzano FM, Bortoloni M, Excoffier L, Dugoujon JM & Ruiz-Linares A. 2010. Contrasting patterns of nuclear and mtDNA diversity in Native American populations. Annals of Human Genetics 74(6): 525-38 [pdf]
  69. Ray N & Excoffier L. 2010. A first step toward inferring levels of long distance dispersal during past expansions. Molecular Ecology Resources 10(5): 902-914 [pdf]
  70. Estoup A, Baird SJE, Ray N, Currat M, Cornuet J-M, Santos F, Beaumont M A & Excoffier L. 2010. Combining genetic, historical and geographic data to reconstruct the dynamics of the bioinvasion of cane toad Bufo marinus. Molecular Ecology Resources 10(5): 886-901 [pdf]
  71. François O, Currat M, Ray N, Han E, Excoffier L & J Novembre. 2010. Principal Component Analysis under population genetic models of range expansion and admixture. Molecular Biology and Evolution 27(6): 1257-1268 [pdf]
  72. Ray N, Wegmann D, Fagundes NJR, Ruiz-Linares A & Excoffier L. 2010. A statistical evaluation of models for the initial settlement of the American continent emphasizes the importance of gene flow with Asia. Molecular Biology and Evolution 27(2): 337–345 [pdf] [suppl. info.]
  73. Ray N & Excoffier L. 2009. Inferring past demography using spatially-explicit population genetic models. Human Biology 81(2-3): 141-157 [pdf]
  74. Janin A, Léna J-P, Ray N, Delacourt C, Allemand P & Joly P. 2009. Assessing landscape connectivity with calibrated cost-distance modelling: predicting common toad distribution in a context of spreading agriculture. Journal of Applied Ecology 46(4): 833-841 [pdf]
  75. Hofer T, Ray N, Wegmann D & Excoffier L. 2009. Large allele frequency differences between human continental groups are more likely to have occurred by drift during range expansions than by selection. Annals of Human Genetics 73: 95-108 [pdf]
  76. Ray N & Ebener S. 2008. AccessMod 3.0: computing geographic coverage and accessibility to health care services using anisotropic movement of patients. International Journal of Health Geographics 7: 63 [pdf]
  77. Excoffier L & Ray N. 2008. Surfing during population expansions promotes genetic revolutions and structuration. Trends in Ecology and Evolution 23(7): 347-351 [pdf]
  78. Excoffier L, Ray N & Currat M. 2008. Where was Paradise? A simulation study of the spread of early modern humans in heterogeneous environments. In Simulations, Genetics and Human Prehistory edited by Matsumura S, Forster P & Renfrew C. (McDonald Institute Monographs.) Cambridge: McDonald Institute for Archaeological Research. Pp. 9-18 [pdf]
  79. Ray N, Currat M & Excoffier L. 2008. Incorporating environmental heterogeneity in spatially-explicit simulations of human genetic diversity. In Simulations, Genetics and Human Prehistory edited by Matsumura S, Forster P & Renfrew C. (McDonald Institute Monographs.) Cambridge: McDonald Institute for Archaeological Research. Pp. 103-117 [pdf]
  80. Currat M, Ray N & Excoffier L. 2008. Genetic simulations of population interactions during past human expansions in Europe. In Simulations, Genetics and Human Prehistory edited by Matsumura S, Forster P & Renfrew C. (McDonald Institute Monographs.) Cambridge: McDonald Institute for Archaeological Research. Pp. 37-47 [pdf]
  81. Wang S, Ray N, Rojas W, Parra M V, Bedoya G, Gallo C, Poletti G, Mazzotti G, Hill K, Hurtado A M, Camrena B, Nicolini H, Klitz W, Barrantes R, Molina J A, Freimer N, Bortolini M C, Salzano F M, Petzl-Erler M L, Tsuneto L T, Dipierri J E, Alfaro E L, Bailliet G, Bianchi N O, Llop E, Rothhammer F, Excoffier L & Ruiz-Linares A. 2008. Geographic patterns of genome admixture in Latin American Mestizos. PLoS Genetics 4(3): e1000037 [pdf]
  82. Neuenschwander S, Largiader CR, Ray N, Currat M, Vonlanthen P & Excoffier L. 2008. Colonization history of the Swiss Rhine basin by the bullhead (Cottus gobio): Inference under a Bayesian spatially explicit framework. Molecular Ecology 17(3): 757-772 [pdf]
  83. Fagundes N J R, Ray N, Beaumont M, Neuenschwander S, Salzano F M, Bonatto S L & Excoffier L. 2008. Ancient lineages and assimilation: A reply to Garrigan and Hammer. Proceedings of the National Academy of Sciences USA 105: E4 [pdf]
  84. Wang S, Jakobsson M, Lewis Jr C M, Ramachandran S, Ray N, Bedoya G, Rojas W, Parra M V, Molina J A, Gallo C, Mazzotti G, Poletti G, Hill K, Hurtado A M, Labuda D, Klitz W, Barrantes R, Cátira Bortolini M, Salzano F M, Petzl-Erler M L, Llop E, Rothhammer F, Excoffier L, Feldman M W, Rosenberg N A & Ruiz-Linares A. 2007. Genetic variation and population structure in Native Americans. PLoS Genetics 3(11): e185 [pdf]
  85. Fagundes N J R, Ray N, Beaumont M, Neuenschwander S, Salzano F M, Bonatto S L & Excoffier L. 2007. Statistical evaluation of alternative models of human evolution. Proceedings of the National Academy of Sciences USA 104: 17614-17619 [pdf] [suppl. info.]
  86. Fontaine M C, Baird S J E, Piry S, Ray N, Tolley K A, Duke S, Birkun A Jr, Ferreira M, Jauniaux T, Llavona A, Öztürk B, Öztürk A A, Ridoux V, Rogan E, Sequeira M, Siebert U, Vikingsson G A, Bouquegneau J-M & Michaux J R. 2007. Rise of oceanographic barriers in continuous populations of a cetacean: the genetic structure of harbour porpoises in Old World waters. BMC Biology 5, 30 [pdf]
  87. Ray N & Burgman M A. 2006. Subjective uncertainties in habitat suitability maps. Ecological Modelling 195: 172-186 [pdf]
  88. Broquet T, Ray N, Petit E, Fryxell J M & Burel F. 2006. Genetic isolation by distance and landscape connectivity in the American marten (Martes americana). Landscape Ecology 21(6): 877-889 [pdf]
  89. Currat M, Excoffier L, Maddison W, Otto SP, Ray N, Whitlock MC & Yeaman S. 2006. Comment on "Ongoing Adaptive Evolution of ASPM, a Brain Size Determinant in Homo sapiens" and "Microcephalin, a Gene Regulating Brain Size, Continues to Evolve Adaptively in Humans". Science 313(5784): 172 [pdf]
  90. Ray N. 2005. PATHMATRIX: a GIS tool to compute effective distances among samples. Molecular Ecology Notes 5(1): 177-180 [pdf]
  91. Ray N, Currat M, Berthier P & Excoffier L. 2005. Recovering the geographic origin of early modern humans by realistic and spatially explicit simulations. Genome Research 15(8): 1161-7 [pdf] [suppl. data]
  92. Hamilton G, Currat M, Ray N, Heckel G, Beaumont M & Excoffier L. 2005. Bayesian estimation of recent migration rates after a spatial expansion. Genetics 170: 409-417 [pdf]
  93. Currat M, Ray N & Excoffier L. 2004. SPLATCHE: a program to simulate genetic diversity taking into account environmental heterogeneity. Molecular Ecology Notes 4(1): 139-142 [pdf]
  94. Ray N, Currat M & Excoffier L. 2003. Intra-deme molecular diversity in spatially expanding populations. Molecular Biology and Evolution 20(1): 76-86 [pdf]
  95. Ray N. 2003. Comment on "Population growth, carrying capacity, and conflict". Current Anthropology 44(1): 77-78 [pdf]
  96. Ray N, Lehmann A & Joly P. 2002. Modeling spatial distribution of amphibian populations: a GIS approach based on habitat matrix permeability. Biodiversity and Conservation 11(12): 2143-2165 [pdf]
  97. Ray N & Adams J M. 2001. A GIS-based vegetation map of the world at the Last Glacial Maximum. 25,000-15,000 BP). Internet Archaeology 11 [pdf] [Data sets download page]


  1. Giuliani G, Lacroix P, Guigoz Y, Bigagli L, Ray N & Lehmann A. 2014. Bringing GEOSS Services into Practice. GIS Open Source Workshop Material. University of Geneva, United Nations Environment Programme, National Research Council of Italy, 189 pages. Available from Google Play Books and from iTunes Store


  1. Ray N. 2003. Modélisation de la démographie des populations humaines préhistoriques à l‘aide de données environnementales et génétiques PhD thesis. Université de Genève, Genève [online thesis].
  2. Ray N. 1999. Etude de la migration des amphibiens et de la connectivité à l‘aide d‘un Système d‘Information Géographique Master thesis (DESSNE). Université de Genève/Université de Lausanne, Genève [pdf]

Other publications

  1. Astsatryan H, Narsisian W, Ghazaryan V, Saribekyan A, Asmaryan Sh, Muradyan V, Guigoz Y, Giuliani G & N Ray. 2012. Toward the Development of an Integrated Spatial Data Infrastructure in Armenia. Proceedings ICT 2012 Conference, pp. 85-92 [pdf]
  2. Joost S, Kalbermatten M & N Ray. 2010. GEOME: Towards an integrated web-based landscape genomics platform. Proceedings of the IUFRO Landscape Ecology Working Group: FOREST LANDSCAPES AND GLOBAL CHANGE - New Frontiers in Management, Conservation and Restoration, IUFRO 2010, Landscape Ecology International Conference, Sept. 21-27, Bragança, Portugal, pp. 668-672 [pdf]
  3. Allenbach K, Burnier J, Lehmann A, Martin P, Ray N and N Wyler. 2010. Cartographie des milieux naturels dans le périmètre Vuache - Sion - Laire. Rapport technique. Projet Franco-valdo-genevois. Cahier No. 13-4. 98 pages
  4. Lehmann A, Giuliani G, Ray N, Allenbach K, Charvat K, Gorgan D, Gvilava M, Bakuradze T. Sözen S, Goksel C & the enviroGRIDS consortium. 2009. The Black Sea Catchment Observation System built on a grid-enabled Spatial Data Infrastructure. INSPIRE, GMES and GEOSS activities, methods and tools towards a single Information Space in Europe for the environment, eds. Charvat, K., Alberts, M. & Horajova, S. Wirelessinfo Tehnologiju attistibas forums, Riga, pp. 42-58 [pdf]
  5. Giuliani, G., Ray, N., Lehmann, A. and K. Charvat. 2009. EnviroGRIDS interoperability guidelines. 92 pages
  6. Gorgan, D., Bacu, V., Ray, N. and A. Maier. 2009. EnviroGRIDS data storage guidelines. 67 pages
  7. Lehmann, A., Ray, N., Kideys, A., Weller, P. and M. Gvilava. 2009. Connecting the enviroGRIDS Black Sea Catchment Observation System to ICZM. Proceedings of the ninth International Conference on the Mediterranean Coastal Environment, MEDCOAST 09, Sochi, Russia, Volume 1, pp. 115-126 [pdf]
  8. Ebener S, Ray N, Black M & El Morjani Z. 2005. Physical Accessibility to health care: from isotropy to anisotropy. GIS Development 9(6): 34-36 [pdf]
  9. Ray, N. and S. Ebener. Physical Accessibility to Health Care and Population coverage Modeling - AccessMod© 3.0. Training and user guide. WHO. 51 pages
  10. Ray N & Adams J. 2002. Present-potential vegetation maps of the world. A GIS-based reconstruction using altitudinal zone [link and download]
  11. Ray, N. 2000. Training Guide for the use of GEO-3 Core Data Sets On-Line and the preparation of spatial data sets. United Nations Institute for Training and Research/Training Programme for the Application of the new Information and Communication Technologies, Geneva
  12. Levine JS, Bobbe T, Ray N, Singh A & RG Witt. 1999. Wildland Fires and the Environment: a Global Synthesis. UNEP-Technical and Assessment Report 99(1) [pdf]


  1. Moser, F., Ebener, S. & Ray, N. 2017. ACCESSMOD 5.0: Supporting Universal Health Coverage by modelling physical accessibility to health care. []
  2. Santiago, H., Ray, N., Ebener, S. 2012. ACCESSMOD 4.0: an ArcGIS extension to model physical accessibility to health care and geographic coverage. [link and download]
  3. Ray N, Currat M, Foll M. & Excoffier L. 2010. SPLATCHE2: a spatially-explicit simulation framework for complex demography, genetic admixture and recombination. [link and download]
  4. Ray, N. 2008. ACCESSMOD 3.0: a GIS tool to compute accessibility to health care based on environmental constraints to movement. [link and download]
  5. Ray, N. 2004. PATHMATRIX: a GIS tool to compute effective distances among samples. [link and download]
  6. Currat, M., Ray, N., Excoffier, L. 2004. SPLATCHE: A program to simulate genetic diversity based on environment heterogeneity. [link and download]
  7. Ray, N. 2002. CORRELATOR. A program for the exploration of correlation among a high number of sampling locations. LGB/University of Geneva
  8. Ray, N. 2002. G.SERVER LINK 1.0. Extension to ArcView 3.x enabling GIS information layers to be extracted and displayed from an IBM DB2-powered G.server. Geotask, Basel, Switzerland
  9. Ray, N. and L. Excoffier. 2002. MANTELN. A program for the computation of correlation and Mantel test on an arbitrary number of matrices. LGB/University of Geneva
  10. Ray, N. 2002. GEODIST. A program to compute matrices of geographic distances. LGB/University of Geneva
  11. Ray, N. 2001. GRID2TEXT and TEXT2GRID. Translator tools between ESRI grid format and regular text files using GRIDIO api. LGB/University of Geneva